To save content items to your account,
please confirm that you agree to abide by our usage policies.
If this is the first time you use this feature, you will be asked to authorise Cambridge Core to connect with your account.
Find out more about saving content to .
To save content items to your Kindle, first ensure no-reply@cambridge.org
is added to your Approved Personal Document E-mail List under your Personal Document Settings
on the Manage Your Content and Devices page of your Amazon account. Then enter the ‘name’ part
of your Kindle email address below.
Find out more about saving to your Kindle.
Note you can select to save to either the @free.kindle.com or @kindle.com variations.
‘@free.kindle.com’ emails are free but can only be saved to your device when it is connected to wi-fi.
‘@kindle.com’ emails can be delivered even when you are not connected to wi-fi, but note that service fees apply.
ABSTRACT The larval chironomid community of the bed sediment surface and the hyporheic interstitial was examined in a gravel stream between September 1984 through August 1985 and between March and June 1993.
Spatial and temporal species turnover between horizontally adjacent sampling sites fluctuated distinctly in all sediment depth layers. The species composition showed a significantly lower spatial turnover in the upper 10 cm of the bed sediments than in deeper layers (P<0.05). Moreover, species abundance patterns of a five-species assemblage implied random assortment (sensu Tokeshi, 1990) and indicated a high probability for a species assembly to be dictated by environmental stochasticity. Neutral models were developed to evaluate the significance of observed overlap values in spatial distribution amongst abundant chironomid species (sensu Schmid, 1993). The spatial organization of a larval chironomid assemblage in the gravel stream Oberer Seebach seemed to be governed by coexistence due to random colonization processes, which reduce the probability of strong competitive interactions. Larval species colonization into open microhabitat patches of interstitial space (47.8 cm3) was rapid with representatives of a species assemblage arriving within less then 24 hours of the start of the colonization experiment. Moreover, possible random movement across and between sediment depth layers of larval chironomid species may promote a rapid colonization in this gravel stream.
INTRODUCTION
The recent emphasis on a non-equilibrium view of communities (Strong et al., 1984), as opposed to an equilibrium one, has drawn attention to the importance of environmental stochasticity, habitat heterogeneity and patchiness in running water ecosystems (Tokeshi, 1994; Schmid, 1993).
By
G. Jacks, Land and Water Resources, Royal Institute of Technology, S-100 44 Stockholm, Sweden,
A. Joelsson, Halland County Board, S-310 86 Halmstad, Sweden,
A.-C. Norrström, Land and Water Resources, Royal Institute of Technology, S-100 44 Stockholm, Sweden,
U. Johansson, Tönnersjöheden Experimental Forest, Swedish University of Agricultural Sciences, S-310 38 Simlångsdalen, sweden
ABSTRACT A small forest catchment in Sweden is drained via a peatland. The mature spruce stand (108 years) was clearcut after which a younger stand (60 years) on the peatland was accidentally stormfelled the next year. The clearcutting causes nitrification in the upland. To maintain the nitrogen reduction function of the peatland, planting on mounds was practiced instead of the conventional fishbone drainage pattern. During the fourth and fifth years after the clearcutting the nitrogen flux from the upland to the peatland was 17 kg ha−1.a while only 7 kg left the peatland. While about 85% of the nitrogen entering the peatland was in the form of nitrate, the fraction of nitrate in the runoff was only 40%. The establishment of new forest stands on peatland soils by planting on mounds gives considerable environmental advantages as compared to conventional drainage. Both fluxes of nutrients and suspended matter are decreased. Spontaneous regeneration of the wetland spruce stand results in loss of about ten years of growth and a younger stand on the wetland than on the upland with the risk of the stormfelling being repeated after the upland forest has been harvested.
INTRODUCTION
Most of the Swedish forest land is drained by a system of ditches. Draining has been done for the purpose of increasing forest production. A peak in the draining operations was seen in the 1930s and it has again increased during the last decades (Löfroth, 1991). A maximum of 12 000 km of drains have been dug yearly, equal to the distance from Stockholm to Vladivostok and back again, much through the same terrain.
By
A. Bothar, Hungarian Danube Research Station, Hungarian Academy of Sciences, H-2131 God, Hungary,
B. Rath, Hungarian Danube Research Station, Hungarian Academy of Sciences, H-2131 God, Hungary
ABSTRACT The 137 km long Szigetköz side-arm system extends on the right side of the river Danube in Hungary and shows the unique characteristics of river, shore, riparian forest, and groundwater ecotones. The investigated three side arms are in different alluvial stages, their surface sediment is variable: sand, sandy mud, mud, mud covered with detritus. The composition, life forms, and cover of aquatic macrophyte stands are good describers of the different side arms. On the surface of the sediment, the habitats are numerous and varied, and linked to the vegetation and open water region. Crustacean communities of these habitats are very diverse, mosaic-form, and are in dynamic connection with each other mainly through the developing stages, and even through certain stenoecious species.
INTRODUCTION
The Szigetköz Danube stretch in Hungary (1848–1806 river km) flows on its own alluvial cone. Fall is small, 3–10 cm km−1. A 137 km long side-arm system extends on the right side of the river, with 22.2 million m3 volume. The whole region is composed of five side-arm systems. Regulation activities began in the last century. The main goal of the engineering has been the assurance of the shipping on the Danube and the flood control. The present situation is the result of the regulation works carried out in 1966–1983 when numbers and heights of closing dams were increased. During low and mean water periods, 90 % of the water discharge flows in the Danube itself. Therefore, quantity and intensity of water supply of the side arms have decreased. The situation of the water table in the Szigetkoz region did not changed drastically in this period.
By
J. Gibert, UNESCO, Division des Sciences Ecologiques, 7 Place de Fontenoy, 75700 Paris, France,
F. Fournier, Université Lyon 1, URA CNRS 1974, Ecologie des Eaux Douces et des Grands Fleuves, Hydrobiologie et Ecologie Souterraines, 43 Bddu 11 novembre 1918, 69622 Villeurbanne cedex, France,
J. Mathieu, UNESCO, Division des Sciences Ecologiques, 7 Place de Fontenoy, 75700 Paris, France
ABSTRACT This paper considers the increased interest in groundwater/surface water ecotones. That clearly appeared within the framework of the UNESCO/MAB and IHP projects on the role of land/inland water ecotones in landscape management and restoration. Groundwater/Surface water ecotones are transition zones, the limits between very contrasted systems. At different space and time scales they provide, favour, filter or stop exchanges and they can also modify interactions between ecosystems. This paper outlines the content and the structure of the book.
BACKGROUND
January 1991 marks the official start of the joint project between the Man and the Biosphere Programme (MAB) and the International Hydrological Programme (IHP) of UNESCO on the role of land/inland water ecotones in landscape management and restoration. The launching of this project was due to the increasing awareness of the important role played by ecotones between terrestrial and aquatic systems in the landscape and in natural resource management (Di Castri et al., 1988; Naiman & Decamps, 1990; Holland & Risser, 1991; Hansen & Di Castri, 1992). They play an essential role as controls for the movement of water and materials throughout the landscape. They are zones where ecological processes are more intense and resources more diversified. The interface favours species dispersal. They are also zones which react quickly to human influences and changes in environmental variables.
Considering the key role of land/inland water ecotones, UNESCO/MAB and IHP launched a collaborative research project with the aim of determining the management options for the conservation and restoration of land/inland-water ecotones through increased understanding of ecological processes. Within this framework, a sub-network was devoted to groundwater/surface water ecotones.
By
G. C. Poole, Center for Streamside Studies, AR-10, University of Washington, Seattle, Washington 98195, USA,
R. J. Naiman, Center for Streamside Studies, AR-10, University of Washington, Seattle, Washington 98195, USA,
J. Pastor, Natural Resources Research Institute, University of Minnesota, Duluth, Minnesota 55811, USA,
J. A. Stanford, Flathead Lake Biological Station, The University of Montana, Poison, Montana 59860, USA
ABSTRACT Ground penetrating RADAR was used in an attempt to map sediment accumulation in active and abandoned beaver {Castor canadensis) ponds in northern Minnesota and to map buried paleochannels of the Flathead River floodplain in Montana. We attempted to map ice thickness, water depth, sediment depth, depth to parent material (bedrock or clay), thickness of soil horizons, organic deposits (peat), frost penetration, and depth to the water table in the beaver ponds. Ground penetrating RADAR successfully located some of the subsurface interfaces between these layers but water saturation and the high clay content of the soils interfered with the ground penetrating RADAR signal while the physical complexity of the subsurface hampered data interpretation. In Montana, paleochannels and water tables were located, but the stony nature of the substrate prevented immediate excavation for verification. In both Montana and Minnesota, success depended strongly on physical characteristics of the sites and specific interfaces. Generally, our efforts were only successful where the physical subsurface interfaces had abrupt, well defined boundaries, and where clay content was low.
INTRODUCTION
Ecological applications of ground penetrating RADAR (GPR) have included classification of soils and remote examination of their structure (Doolittle, 1982; Ulriksen, 1982).
By
M. Trémolières, Laboratoire de Botanique et Ecologie Végétale, CEREG URA 95 CNRS, Institut de Botanique, 28 rue Goethe, F-67083 Strasbourg cedex, France,
D. Correll, Smithsonian Environmental Reasearch Center, P.O. Box 28, Edgewater, Maryland 21037, USA,
J. Olah, Fisheries Research Institute, H-5541 Szarvas, Hungary
The overall goal of this research on riparian zones is to understand how these habitats function and what factors control their functions. The water quality factors of greatest interest are nutrients and toxic materials. The original publications which reported on these buffering functions of riparian forest were Gilliam et al. (1974), and Gambrell et al (1975). These were followed by a series of more focused reports (Lowrance et al, 1984a, 1984b; Peterjohn & Correll, 1984; 1986; Labroue & Pinay, 1986; Pinay & Labroue 1986; Schnabel, 1986; Pinay & Decamps, 1988; Correll & Weller, 1989; Sanchez-Perez et al, 1991, 1993).
Comparative data are needed to understand the role of plant communities in different riparian systems and interaction processes between compartments (Fig. 1). Such questions need to be addressed to ascertain the relative effectiveness of grass, herbs and trees; the width of vegetation needed for effectiveness; the importance of primary production, plant diversity, age structure, depth of root/rhizosphere zones; and how the surface plant community interacts with below-ground microbial communities. These systems should be characterized in respect to their hydrology, geomorphology, biogeochemistry, below-ground conditions, and hyporheic zones. More similarity needs to be developed in approaches and methods in order to produce more comparable data in the future.
GENERAL QUESTIONS FOR FUTURE RESEARCH
a) What are the capacities of these systems for processing nutrients and toxins?
b) Are these systems self-sustaining? For how long?
c) What are the principal mechanisms of water quality affects and what controls their rates?
d) What is the importance of biodiversity in controlling the efficiency of riparian zone processing of man-made chemicals?
By
PH. Vervier, CERR–CNRS, 29 rue Jeanne Marvig, 31055 Toulouse, France,
M. H. Valett, Department of Biology, University of New Mexico, Albuquerque, New Mexico 97131, USA,
C. C. Hakenkamp, Department of Zoology, University of Maryland at College Park, 1200 Zoology-Psychology Building, College Park, MA 20742–4415, USA,
M.-J. Dole-Olivier, HBES, URA 1974, UCB Lyon 1, 43 Bddu 11 novembre 1918, 69622 Villeurbanne cedex, France
The aim of this work was to examine the contribution of the groundwater/surface water (GW/SW) ecotone concept towards a better understanding of ecosystem functioning in running water systems. It became clear that a consideration of the differences and similarities between the GW/SW ecotone and the hyporheic zone (HZ) concepts would be helpful. Four major parts could be pointed out. The first part focuses on a comparison between the GW/SW ecotone concepts. The second part emphasizes how the interaction zone between GW and SW influences ecosystem functioning in the two adjacent systems. The third part suggests important topics for future research, and finally recommendations for UNESCO and for managers of river ecosystems are made in the fourth part.
ECOTONE VERSUS HYPORHEIC ZONE
Origin of terms
The hyporheic zone was described in Germany 30 years ago (Orghidan, 1959; Schwoerbel, 1964) but has only recently flourished in the vocabulary of North American stream ecologists (e.g. Williams, 1984; Stanford & Ward, 1988; Triska et al, 1989). Though the term originally associated with a strong biological bias, it is now used in a more general manner to describe the deep sediments of stream beds where GW/SW exchange (see Valett et al, 1993; Hakenkamp et al, 1993).
The ecotone concept is much older since Clements (1904) used this term to describe contact zones between adjacent communities. The ecotone concept also used by Leopold (1933) and Odum (1971), had an initial organismal or community emphasis.
By
A.M.J. Meijerink, International Institute for Aerospace Survey, and Earth Sciences (ITC), Enschede 7500 AA, The Netherlands,
W. van Wijngaarden, International Institute for Aerospace Survey, and Earth Sciences (ITC), Enschede 7500 AA, The Netherlands
ABSTRACT The Amboseli ecosystem consists of the basement plains, the lacustrine saline plains with fresh water swamps and the volcanic slopes of the Kilimanjaro. The area is well known for its large and varied population of wild herbivores, supporting a large tourist industry. The two major fresh spring zones sustaining some 20 km2 of swamps and an additional 16 km2 of wet areas chiefly covered by grass, belong to the relatively shallow part of a regional groundwater flow system in the volcanic complex. A 7.5 m rise of the groundwater table since the 1960s and the recent expansion of one of the swamps has caused concern. The rise cannot be explained by the available rainfall data and may be related to more frequent runoff, because of overgrazing, in the catchment feeding episodically the seasonal Lake Amboseli. In addition, tectonic movements may have increased the outflow of the the deeper diffuse part of the flow system.
The dynamics of the swamps have been studied. Available data suggest that no major changes are likely to affect the swamps in the near future. A good proportion of the annual recharge of the springs, about 14 % of the estimated rainfall, takes place in the Tanzanian part.
INTRODUCTION
The Amboseli ecosystem can be characterized as a semiarid savanna environment, which shows considerable spatial and temporal variation in resources; climate, soils, vegetation and hydrology. This has an important bearing on the structure and functioning of the ecosystem and has important consequences for the management. The area is well known for its large and varied population of wild and domestic herbivores, upon which a large tourist industry is based.
ABSTRACT Vertical and temporal distribution of paniculate organic matter (POM) in river bed sediments was observed in two different stations in the Morava River. Samples of river sediments from 0–70 cm depths were taken by inserting a steel cylinder with a special grab. The main type of organic matter found in bed sediments was fine particulate organic matter (FPOM < 1 mm). The organic matter had a variable distribution in the hyporheic zone. Generally, the concentration of organic matter was higher in the top 0–40 cm of sediment. The annual mean storage of buried organic matter (0–70 cm depth) was 5352.13 g AFDW.rn−3. The exchange of organic matter between surface sediments and deep river bed sediments is influenced mainly by flooding activity. The maximum storage of organic matter in the hyporheic zone appears in the spring when floods are decreasing.
INTRODUCTION
Allochthonous organic matter which is imported in running waters at different times and in different ways (Boling et al, 1975; Cummins, 1974; Moser, 1991) is an important source of energy for sediment fauna (Fisher & Likens, 1973; Cummins, 1974). Fresh imported allochthonous organic matter is not available directly for benthic consumers and must first be processed by the microbial communities, especially fungi and bacteria (Rossi & Fano, 1979; Arsuffi & Suberkropp, 1985).
An exchange of water exists between the surface and hyporheic zone (Grimm & Fisher, 1984; Thibodeaux & Boyle, 1987; White et al, 1987). In these exchange processes, particulate and dissolved organic material are transported into the bed sediments (Welton, 1980; Carling & McMahon, 1987; Mayack et al, 1989; Metzler & Smock, 1990; Smock, 1990; Bretschko & Moser, 1993; Sear, 1993).
By
F. Malard, Université Lyon 1, URA CNRS 1974, Ecologie des Eaux Douces et des Grands Fleuves, Hydrobiologie et Ecologie Souterraines, 43 Bd du 11 novembre 1918, 69622 Villeurbanne cedex, France,
M.-J. Turquin, Université Lyon 1, URA CNRS 1974, Ecologie des Eaux Douces et des Grands Fleuves, Hydrobiologie et Ecologie Souterraines, 43 Bd du 11 novembre 1918, 69622 Villeurbanne cedex, France,
G. Magniez, Laboratoire de Biologie Animate et Générate, Université de Bourgogne, 6 Bd Gabriel, 21000 Dijon, France
ABSTRACT Faunal sampling was carried out in 1990–91 on three springs of a karstic system located at 15 km north-east of the city of Dijon (France) to investigate the filter effect of these outlets on the drifting population of the hypogean amphipod Niphargus virei. Results showed that, in places where limestones outcropped, the karstic spring ecotone did not modify the structure of the drifting population. In that case, faunal sampling of the springs during the floods was an effective method to study the dynamics of the population living in the aquifer. However, this method could not be applied in places where deep karstic ground-water circulated through water-saturated superficial deposits prior to emerge at the land surface. Indeed, in that case, the size frequency histograms of the drifting population of N. virei were characterised by abnormally low percentages of small size individuals and/or to a lesser extent by low percentages of large size individuals. These changes in the population structure were probably caused by two types of filters exerted by the karstic spring ecotone: 1) a mechanical filter which corresponded to the change of matrix between the deep karstified limestone aquifer and the shallow aquifer; 2) a biological filter probably due to the predation exerted by benthic invertebrates which have colonised the shallow aquifer.
INTRODUCTION
Surface water/groundwater ecotones (i.e. interaction zones occurring between surface water and groundwater systems) regulate the flow of matter, energy, information and organisms between two contrasted ecological systems (UNESCO, 1980; Pennak & Ward, 1986; Vanek, 1987; Ford & Naiman, 1989; Gibert, 1991a; Sharley, 1994).
By
J. Simons, Environmental Protection Agency, 401 M Street SW, Washington DC 20460, USA,
J. Notenboom, RIVMLaboratory of Ecotoxicology, P.O. Box 1, Bilthoven 3720 BA, The Netherelands
The scientific community is increasingly interested in becoming involved in management of groundwater resources and of related surface water and riparian ecosystems impacted by groundwater/surface water interactions. Researchers want more control over how the results of their work are applied and they want to be more involved in the important task of environmental protection. They also want to have more influence on research funding. In fact, many scientists feel their involvement is necessary for their work to continue receiving support and funding. This new involvement of researchers is welcomed by enlightened water resources managers. Managers are under increasing pressure to balance protection policies with an array of other needs. They realize that the most effective management must be based on the best scientific information. Because of these factors, scientists are more frequently devoting a portion of their programmes to management issues (Stanford & Simons, 1992). However, groundwater management is complex, dealing with scientific as well as social, economic and political issues. There are many problems that make solutions difficult, and progress is often slow. In spite of this, scientists and managers must continue to push for a more co-ordinated approach because protection of our vital groundwater resources needs the involvement of everyone who can help.
GOAL STATEMENT
To base the management of groundwater resources on the best available scientific information in order to achieve maximum ecological benefits. By doing this, groundwater will also be protection as a safe source of drinking water.
A list of actions is needed in order to achieve this goal, and how these actions could be implemented. These actions are really the basis for developing a strategic plan of action.
By
P. MARMONIER, Université de Savoie, GRETI, 73376 he Bourget du Lac, France,
J.V. WARD, Department of Biology, Colorado State University, Fort Collins, Colorado 80523, USA,
D.L. DANIELOPOL, Limnological Institute, Austrian Academy of Sciences, A-5310 Mondsee, Austria
Biodiversity is the object of a large international programme of the IUBS-SCOPE-UNESCO and is important for many scientific, economic and ethic reasons (Solbrig, 1992). In groundwater, the study of biodiversity can be considered as a promising research field (Marmonier et al, 1993). Five major topics were discussed:
the importance of groundwater/surface water ecotone fauna in the estimation of global diversity;
the local and regional biodiversity;
factors promoting biodiversity;
biodiversity in gradients and impact of disturbances;
how to preserve biodiversity.
IMPORTANCE OF GROUNDWATER/SURFACE WATER ECOTONE FAUNA IN THE ESTIMATION OF GLOBAL DIVERSITY
Groundwater fauna is largely ignored in the calculation of global biodiversity. The number of species of tropical rain forests is estimated to be 10 or 20 times higher than those known to science (Cairns, 1988). Groundwater diversity is less well known than the diversity of surficial species in tropical forests globally: the total biodiversity of groundwater fauna is certainly higher than current estimations (in the Stygofauna Mundi, for example – Botosaneanu, 1986) and these organisms may represent an important part of the global biodiversity.
This is greatly important if groundwater fauna play an active role in groundwater system functioning (Fig. 1). It is especially true for microbes (microfauna, bacteria, fungi) which are still more or less unknown and may play an important role in the bank filtration of large rivers (the River Rhine for example is rather well studied from this point of view).
Groundwater/surface water interactions are often characterised by hydrological and hydrochemical processes which result in complex abiotic and biotic gradients. Therefore a study was made of the hydrochemistry and ecohydrology of a groundwater discharge area in a low coastal area, lying in front of a higher sandy recharge area in the southwest of the Netherlands.
A hydrochemical facies analysis was applied to identify and map the major factors accounting for variations in hydrochemical processes. Research was concentrated in a relatively small fresh water upwelling zone in the discharge area, to obtain insight into the actual interrelationship between groundwater, surface water, aquatic ecology and human influence. During 1992 surface water samples and groundwater samples were taken along four regional transects. In the upwelling zone, sampling of shallow groundwater and surface water was combined with detailed mapping of aquatic vegetation. In total, 43 species were found and over 275 water samples were analysed in the laboratory. The two main water systems, the Brabantse Wai hydrosome and the polder hydrosome, are described by seven chemical watertypes, redox potential index, eutrophication potential index and calcite saturation index. The regionally derived maps of ground and surface water quality correspond fairly accurately. In the fresh water upwelling zone four types of aquatic vegetation (noda) have been discerned. Each nodum has distinct ecological amplitudes for alkalinity, pH and salinity of the surface water. These do not match general indicative values for the Netherlands, as described in literature. The hydrochemical characteristics of the ground and surface water in the upwelling zone deviate to a large extent. This points to an increasing contribution of polluted groundwater from a shallow groundwater flow system.…
By
J. V. Ward, Department of Biology, Colorado State University, Fort Collins, Colorado 80523, USA,
N. J. Voelz, Department of Biology, St. Cloud State University, St. Cloud, Minnesota 56301, USA
ABSTRACT Interstitial animals were collected during spring, summer, and autumn from eight sampling sites along a Rocky Mountain river to examine small scale patterns of diversity, abundance, and faunal composition across the groundwater/surface water ecotone. At each site samples were taken from benthic habitats (superficial bed sediments), hyporheic habitats (underflow 30 cm below the bed surface), and phreatic habitats (30 cm below the water table of adjacent alluvial bars). Total taxa decreased markedly along the epigeanhypogean gradient. Crustaceans and insects (the majority of animals in all three habitats) progressively increased (Crustacea) or progressively declined (Insecta) in relative abundance along the epigean-hypogean gradient. Most of the 142 common taxa exhibited one of four types of distribution patterns along the gradient (hypogean, epigean, transitional, eurytopic). This study demonstrated a marked faunal gradient on a scale of meters across the groundwater/surface water ecotone. Faunal similarity coefficients (e.g., Jaccard) and detrended correspondence analysis of faunal distributions may provide comparative measures of ecotone permeability between physically contiguous interstitial habitats.
INTRODUCTION
Riverine sediments form habitat patches of various sizes and complexity, reflecting the interactions of fluvial dynamics and biogeochemical features (Schumm, 1977; Richards, 1982; Rust, 1982). The distribution patterns of animals inhabiting porous alluvia reflect gradients in environmental conditions that occur at a variety of spatial scales (reviewed by Gibert et al., 1994; Ward & Palmer, 1994). Interstitial animals residing in the water-filled spaces between substrate particles are distributed along a gradient of habitat conditions from superficial bed sediments (benthic biotope), to deeper sediments beneath the channel (hyporheic biotope=underflow), to alluvial deposits situated some distance laterally from the active channel (phreatic biotope).
By
V. Vanek, Université Lyon 1, URA CNRS1974, Ecologie des Eaux Douces et des Grands Fleuves, Hydrobiologie et Ecologie; Souterraines, 43 Bddu 11 novembre 1918, 69622 Villeurbanne cedex, France; VBB VIAK Consulting Engineers, Geijersgatan 8, S-216 18 Malmö, Sweden
ABSTRACT Groundwater-surface water ecotones control hydraulic exchange between groundwater and surface water and, together with land-water ecotones, serve as a temporary or permanent sink of catchment-derived inorganic and organic matter. These factors and the chemistry of water entering an ecotone, determine the ecotone's effect on water quality One of the problems when studying ecotones is their high heterogeneity in space and time. The heterogeneity is caused by several thousand years of human impact, which has been exacerbated during the last century, and by local geology, surface water effects and various ecotonerelated processes (capillary-fringe effect, evapotranspiration, anti-clogging, clogging). The heterogeneity is measurable and at least partly predictable, and should be considered when preparing sampling strategies and evaluating data. Ecotones often function as filters of nutrients and particulate matter, thus improving the quality of water passing through them. The adsorption or buffering capacity of the ecotone, however, may be exhausted which may result in the enrichment of water by organic matter, dissolved metals etc. This ‘malfunctioning’ occurs often naturally as a result of decomposition of accumulated organic matter, but may also be due to the excessive leaching of fertilizers or other substances from upstream areas. Losses from catchments may be diminished by increasing the area and improving the function of the ecotones. The ecotone-oriented measures, however, must be combined with modified land use of the whole catchments if we want to use the landscape in a sustainable way.
By
V. Vanek, Université Lyon 1, URA CNRS 1974, Ecologie des Eaux Douces et des Grands Fleuves, Hydrobiologie et Ecologie Souterraines, 43 Bd du 11 novembre 1918, 69622 Villeurbanne cedex, France; Present address: VBB VIAK Consulting Engineers, Geijersgatan 8, S-216 18 Malmö, Sweden,
C. Thirriot, Institut de Mécanique des Fluides, URA 0005 CNRS, Allée du Professeur Camille Soula, 31400 Toulouse, France,
A.M.J. Meijerink, International Institute for, Aerospace Survey ard Earth Sciences (ITC), Enschede, 7500 AA, The Netherlands,
G. Jacks, Land and Water Resources, Royal Institute of Technology, S-10044 Stockholm, Sweden
Modelling the flows across the groundwater/surface water ecotone often concentrates around the following questions: (1) why should we model, (2) what kind of fluxes and scales should be considered in our models, (3) possibilities and difficulties when using models developed for other purposes, (4) communication gaps and future research needs.
(1) The modelling of ecotone-related fluxes may (and should) be used in the following cases:
– to test assumptions and hypotheses
– to improve our understanding and insight into the processes
– to design new experiments and to optimize sampling strategies
– to generalize and synthesize the results
– to develop scenarios and make predictions.
The development of scenarios is felt as one of most important tasks. The ecotones are subjected to the increasing pressure of various human activities (river flow regulation, groundwater extraction, irrigation, non-point pollution etc.), and we need more efficient tools to predict possible impacts of these activities and to improve the management practices. The situation is particularly critical in the third world where hundreds of millions of people live on the surface/groundwater ecotones, depending on them and affecting their function.
(2) The groundwater/surface water ecotones convey numerous fluxes – of water, heat, solutes, particulate matter, genetical information etc. (Gibert et al, 1990). All these fluxes occur on different spatial and temporal scales (Franzle & Kluge, in prep.).
By
D.L. Danielopol, Limnological Institute, Austrian Academy of Sciences, A-5310 Mondsee, Austria,
R. Rouch, Centre de Recherches Souterraines et Edaphiques, Laboratoire Souterrain, CNRS, F-09200 Moulis, France,
P. Pospisil, Institute of Zoology, University of Vienna, Althanstrasse 14, A-1090 Vienna, Austria,
P. Torreiter, Institute of Zoology, University of Vienna, Althanstrasse 14, A-1090 Vienna, Austria,
F. Möszlacher, Institute of Zoology, University of Vienna, Althanstrasse 14, A-1090 Vienna, Austria
ABSTRACT In order to better understand the structure of the surface-groundwater organismal assemblages (SGOA), the concept of global aquifer/river system (GARS) of Castany is recalled. The pattern of SGOAs emerges within different space and time scales. Examples of SGOAs and of the processes which form them are presented using case studies from the alluvial sediment fauna of the rivers Rhone, the Danube at Vienna, and the Lachein brook, in southern France. The structure of these assemblages depends on the boundary conditions of the groundwater system, mainly the hydrologic dynamics, and on the initiating conditions offered by the ecological tolerances of each species which contributes in the assemblages. The possible role of the macroorganisms in the functioning of the groundwater ecosystem is discussed. Their role appears less important than was thought earlier. Finally we emphasize the necessity of protecting the diversity of ecotonal assemblages. Scientific, cultural and practical arguments are presented.
INTRODUCTION
The review deals with studies on surface-groundwater organismic assemblages (SGOA) in unconsolidated, porous media, mainly alluvial sediments. These animal assemblages are formed by hypogean dwelling organisms (Hy) and epigean ones (Epi) which live permanently or temporarily in such subsurface habitats. At a first approximation, the SGOA definition agrees with the concept of ecotone, sensu Odum (1971, p. 157): ‘a transition between two or more diverse communities’. The study of such ecotonal animal assemblages represents one of the core aspects of groundwater ecology (GW).