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Published online by Cambridge University Press: 26 September 2025
The family Glassiidae from Silurian and Devonian strata exhibits barrel-shaped and medially directed spiralia as primary characteristics. Membership in this family is small with five known genera, of which only two possess recognized spiralia. This paper aims to investigate the two Devonian glassiides: Karbous Havlíček, 1985 and Trigonatrypa Havlíček, 1990. In mature specimens, the type species K. aperinus Havlíček, 1985 and the species K. hassiacus (Siehl, 1962) show dorsally directed spiralia. Karbous is here placed in the family Karpinskiidae based on similar internal characteristics. The type species of Trigonatrypa was not available for study. A closely related species, T. baucis (Barrande, 1847), was sectioned and found to possess medially directed spiralia. The type species is likely to have the same kind of spiralia. Therefore, the case for placing Trigonatrypa in the Glassiidae is strengthened.
Handling Editor: Colin Sproat